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Wolf History, Conservation, Ecology and Behavior
Abridged version available in RESOURCES
Article available in WOLF WARS
Abstract available in ABSTRACTS

Wolves Bibliography-S
Sablin, M.V. & G.A. Khlopachev. The Earliest Ice Age Dogs: Evidence from Eliseevichi 1. Current Anthropology, 43/5, December 2002, 795 (5pp).
Saetre, P., et al. From Wild Wolf to Domestic Dog: Gene Expression Changes in the Brain. Molecular Brain Research, 126/2, July 2004, 198 (9pp).
Sage, Jr., R.W. Wolves in the Adirondacks? Perspectives from the Heart of the Adirondack Park. In (Sharpe, V.A. et al., eds.) Wolves and Human Communities: Biology, Politics, and Ethics, 2001, 39-46.
Sahr, D.P. & F.F. Knowlton. Evaluation of Tranquilizer Trap Devices (TTDs) for Foothold Traps Used to Capture Gray Wolves. Wildlife Society Bulletin, 28/3, Fall 2000, 597.
Sandin, Jo. Wolves Go On the Attack: Dogs' Deaths Hit Record Level. Milwaukee Journal-Sentinel, 30 September 2001.
Sands, J.L. & S. Creel. Social Dominance, Aggression and Fecal Glucocorticoid Levels in a Wild Population of Wolves, Canis lupus. Animal Behaviour, 2004 (in press).
Savolainen, P. Genetic Evidence for an East Asian Origin of Domestic Dogs. Science, 298, 22 November 2002, 1610 (4pp).
Sawyer, E.J. American Wolves. Western Sportsman, 5/3, August 1940, 28.
Sax, J.L. Legal and Policy Challenges of Environmental Restoration. In (Sharpe, V.A. et al., eds.) Wolves and Human Communities: Biology, Politics, and Ethics, 2001, 109-109-127.
Schadler, C.S. Reintroduction: Inspired Policy or Poor Conservation? In (Sharpe, V.A. et al., eds.) Wolves and Human Communities: Biology, Politics, and Ethics, 2001, 161-173.
Schadler, C.S. & J. Hammill. Status of the Gray Wolf in Michigan: A Naturally Recovering Population. In (Fascione, N & M. Cecil, comps.) Wolves of America Conference Proceedings, November 1996, 104-110.
Schaefer, C.L. Spatial and Temporal Variation in Wintering Elk Abundance and Composition, and Wolf Response on Yellowstone's Northern Range. Unpublished, thesis, Michigan Technicological University, 2000.
Schantz, V.S. An Unusual Specimen of Red Wolf. Journal of Mammalogy, 17/4, November 1936, 415.
Schauffler, B.F. Wolving -- With Complications. Outing, 60/2, May 1917, 173-176.
Schlickeisen, R. Overcoming Cultural Barriers to Wolf Reintroduction. In (Sharpe, V.A. et al., eds.) Wolves and Human Communities: Biology, Politics, and Ethics, 2001, 61-74.
Schmidt, P.A. & L.D. Mech. Wolf Pack Size and Food Acquisition. American Naturalist, 150/4, October 1997, 513-517.
Schmidt, K.P. & J.R. Gunson. Evaluation of Wolf-Ungulate Predation Near Nordegg, Alberta: Second Year progress Report, 1984-85. Alberta Energy and Natural Resources, 1985 (53pp).
Schneider, S.M. Wasp's Nests and Wolf Packs: A Note on Extending Behavior Setting Theory to Non-Human Groups. Journal of Environmental Psychology, 10/4, 1990, 371-376.
Schropfer, R. & A. Rhode. The Integration of a Young Female in a Male Pack of European Wolves Canis lupus. International Journal of Mammalian Biology, 62, 1997, 209-213.
Schullery, P. & L. Whittlesey. The Documentary Record of Wolves and Related Wildlife Species in the Yellowstone National Park Area Prior to 1992. In (Varley, J.D. & W.G. Brewster, eds.) Wolves for Yellowstone? A Report to the United States Congress, Vol. 4: Research and Analysis, Yellowstone National Park (WY): National Park Service, 1992, 1-267.
Science Weekly. Dogs and Wolves. 19/15, 11 April 2003, 1 (12pp).
Scott, B.M.V. & D.M. Shackleton. Food Habits of Two Vancouver Island Wolf Packs: A Preliminary Study. Canadian Journal of Zoology, 58, 1980, 1203-1207.
Scott, J.P. The Social Behavior of Dogs and Wolves: An Illustration of Sociobiological Systematics. Annals of the New York Academy of Sciences, 51, 1950, 1009-1021.
Scott, P.A. et al. Aggressive Behavior by Wolves Toward Humans. Journal of Mammalogy, 66, 1985, 807-809.
Seal, U.S. et al. Blood Analyses of Wolf Pups and Their Ecological and Metabolic Interpretation. Journal of Mammalogy, 56, 1975, 64-75.
Sears, H.J. A Landscape-Based Assessment of Canis Morphology, Ecology and Conservation in Southeastern Ontario. MSc thesis, University of Waterloo (Waterloo, ON), 1999 (132pp).
Sen, S. Wolves Blamed for Ecological Imbalances. Alternatives, 21/2, April 1995, 10.
Servin, J. Some Aspects of the Social Behaviour of the Mexican Wolf (Canis lupus baileyi) in Captivity. Acta Zoologica Mexicana Nueva Serie, 45, 1991, 1-28.
Seton, E.T. The Habits of Wolves. American Magazine, 64/6, October 1907, 636-645.
Shahi, P.S. Status of the Grey Wolf (Canis lupus pallipes) in India: A Preliminary Survey. Journal of the Bombay Natural History Society, 79/3, 1982, 493-502.
Sheffy, L.F. The Lobo as a Factor in the Cattle Industry. The Cattleman, 15/10, 1929, 94-99.
Shelton, P.C. Ecological Studies of Beavers, Wolves and Moose in Isle Royale National Park, Michigan. Ph.D. Thesis, Purdue University, 1966 (308pp).
Shepard, E.E. Wolves Kill Foxes. Hunting and Fishing in Canada, 6/11, 1940, 37.
Shields, W.M. Genetic Considerations in the Management of the Wolf and Other Large Vertebrates: An Alternative View. in (Carbyn, L.N., ed.) Wolves in Canada and Alaska, Canadian Wildlife Service, Report Series 45, 1983, 90-92.
Shivik, J.A. et al. Current Development and Future Directions in Non-Lethal Wolf Predation Management. World Wolf Congress 2003.
Shivik, J.A. et al. Electronic Aversive Conditioning for Managing Wolf Depredation. Proceedings of the Vertebrate Pest Conference, 20, 2002, 227-231.
Shivik, J.A. et al. Nonlethal Techniques for Managing Predation: Primary and Secondary Repellants. Conservation Biology, 17, 2003, 1531-1538.
Shivik, J. The Other Tools of Wolf Management. WOLF!, 11, 2001, 3-7.
Shogren, E. Saving the Wolves Comes With A Price. Calgary (Alberta) Herald, 15 July 2002, A3.
Sidorovich, V.E. et al. Wolf (Canis lupus) Numbers, Diet and Damage to Livestock in Relation to Hunting and Ungulate Abundance in Northeastern Belarus, 1990-2000. Wildlife Biology, 9/2, June 2003, 103-111.
Sillero-Zubiri et al. Bale Mountains Rodent Communities and Their Relevance to the Ethiopian Wolf (Canis simensis). African Journal of Ecology, 33/4, 1995, 301-320.
Sillero-Zubiri, C. & D. Gottelli. Diet and Feeding Behavior of Ethiopian Wolves (Canis simensis). Journal of Mammalogy, 76/2, May 1995, 531.
Sillero-Zubiri, C. Field Immobilization of Ethiopian Wolves (Canis simensis). Journal of Wildlife Diseases, 32/1, 1996, 147-151.
Sillero-Zubiri, C. et al. A Hypothesis for Breeding Synchrony in Ethiopian Wolves (Canis simensis). Journal of Mammalogy, 79/3, 1998, 853-858.
Sillero-Zubiri, C. et al. Male Philopatry, Extra Pack Copulations and Inbreeding Avoidance in Ethiopian Wolves (Canis simensis). Behavioral Ecologfy and Sociobiology, 38/5, 1996, 331-340.
Sillero-Zubiri, C. et al. Rabies and Mortality in Ethiopian Wolves (Canis simensis). Journal of Wildlife Diseases, 32/1, 1996, 80-86.
Sillero-Zubiri, C. & D.W. Mcdonald. Scent-Marking and Territorial Behaviour of Ethiopian Wolves (Canis simensis). Journal of Zoology, 245/3, 1998, 351-261.
Sillero-Zubiri, C. & D. Gotelli. Spatial Organization in the Ethiopian Wolf Canis simensis: Large Packs and Small Stable Home Ranges. Journal of Zoology, 237, 1995, 65-81.
Silver, J. Hunting the Den of the Timber Wolf. The Northern Sportsman, 3/12, April 1933, 3-4, 14.
Singer, F.C. & J. Dalle-Molle. The Denali Ungulate-Predator System. Alces, 21, 1985, 339-358.
Singer, F.J. Some Predictions Concering a Wolf Recovery into Yellowstone National Park: How Wolf Recovery MayAffect Park Visitors, Ungulates, and Other Predators. Transactions of the North American Wildlife and Natural Resources Conference, 56, 1991, 567-583.
Singer, F. J. Status and History of the Timber Wolves in Glacier National Park, Montana. In (Klinghammer, E., ed.). The Behavior and Ecology of Wolves, 1979, 19-42.
Singer, F.J. The Ungulate Prey Base for Wolves in Yellowstone National Park. In (Keiter, R.B. & M.S. Boyce, eds) The Greater Yellowstone Ecosystem (New Haven: Yale Univ. Press, 1991), 323.
Singleton, P. H.. 1995. Winter Habitat Selection by Wolves in the North Fork of the Flathead River Basin, Montana and British Columbia. M.Sc. Thesis, Univeristy of Montana, 1995 (116 pp).
Skeel, M.A. & L.N. Carbyn. The Morphological Relationships of Gray Wolves (Canis lupus) in National Parks of Central Canada. Canadian Journal of Zoology, 55, 1977, 737-747.
Skogen, K. Who's Afraid of the Big Bad Wolf? Young People's Response to the Conflicts Over Large Carnivores in Eastern Norway. Rural Sociology, 66/2, June 2001, 203.
Skogen, K. & O. Krange. A Wolf at the Gate: The Anti-Carnivore Alliance and the Symbolic Construction of Community. Sociologia Ruralis, 43/3, July 2003, 309 (17pp).
Skoog, R.O. Results of Alaska's Attempts to Increase Prey by Controlling Wolves. Acta. Zool. Fennica, 174, 1983, 245-247.
Skurzynski, G. Wolves: Home Free, But for How Long? National Geographic World, 273, May 1998, 7-11.
Sleeman, Sir W.H. Wolves Nurturing Children in Their Dens. The Zoologist (3d series), 12, 1888, 87-98.
Slobokin, L.B. How To Be A Predator. American Zoologist, 8, 1968, 43-81.
Slobokin, L.B. Prudent Predation Does Not Require Group Selection. American Naturalist, 108, 1974, 665-678.
Smart, G.F. A Wolf Pack Howls in His Back Yard. American Magazine, 111/2, February 1931, 81, 129.
Smirnov, M.N. & V.V. Shurygin. Winter Coloration of Hair-Covering of Wolves. Tuva, Byulleten Moskovskogo Obshchestva Ispytatelei Prirody Otdel Diologicheskii, 99/1, 1991, 14-16.
Smith, C. Report Puts Park Wolves in Dire Peril. The Toronto Star, 13 January 2001.
Smith, C. Why We Should Let Wolves Be. The Toronto Star, 6 January 2001.
Smith, D.T. et al. Is Incest Common in Gray Wolf Packs? Behavioral Ecology, 8/4, 1997, 384-391.
Smith, D. W. et al. Killing of Bison (Bison bison) calf, by a wolf (Canis lupus), and four coyotes (Canis latrans), in Yellowstone
 National Park. Canadian Field-Naturalist. 115/2, 2001, 343-345.
Smith, D.W. & D.R. Stahler. Management of Habituated Wolves in Yellowstone National Park. Yellowstone National Park: Yellowstone Center for Resources, National Park Center, 2003.
Smith, D.W. et al. Managing Wolves and Humans in Lamar Valley: A Final Report on the Druid Road Project. Yellowstone National Park Report, 2000 (5pp).
Smith, D.W. et al., Managing Wolves and Humans in Lamar Valley: A Final Report on the Druid Road Project 2001. Yellowstone National Park Report, 2001 (7pp)
Smith, D.W. et al. Managing Wolves and Humans in Lamar Valley: A Final Report on the Druid Road Project, 2002. Yellowstone National Park Report, 2002 (9pp)
Smith, D.W. & M.K. Phillips. Northern Rocky Mountain Wolf (Canis lupus nubilus). In (Reading, R.P. & B. Miller (eds.), Endangered Animals: A Reference Guide to Conflicting Issues, Greenwood Press, Westport, CT., 2000, 219-233.
Smith, D.W. Wildlife Art: Does It Make A Difference for Wolves? Wildlife Art. 20/6, 2000, 102-105.
Smith, D.W. Wolf #7: The Passing of A Matriarch. Yellowstone Science 10, 2001,18-19.
Smith, D.W. et al. Wolf-Bison Interactions in Yellowstone National Park. Journal of Mammalogy, 81/4, November 2000, 1128.
Smith, D.W. Wolves and Human Communities: Biology, Politics and Ethics. Journal of Mammalogy, 83/3, August 2002.
Smith, D. W. et al. Wolves in the Greater Yellowstone Ecosystem: Restoration of a Top Carnivore in a Complex Management Environment. In Clark, T. et al. (eds.), Carnivores in Ecosystems, 1999, 103-125.
Smith, D.W. The Wolves of Yellowstone. Southeastern Wildlife Magazine, 2000.
Smith, D.W. et al. Yellowstone After Wolves. Bioscience, 53/4, April 2003, 330-340.
Smith, D.W. et al. Yellowstone Wolf Project: Annual Report 2002. National Park Service, Yellowstone Center for Resources, Yellowstone National Park, 2003.
Smith, D.W. & D.S. Guernsey. Yellowstone Wolf Project: Annual Report 2001. National Park Service, Yellowstone Center for Resources, Yellowstone National Park, 2001.
Smith, D.W. & D.S. Guernsey. Yellowstone Wolf Project: Annual Report 2000. National Park Service, Yellowstone Center for Resources, Yellowstone National Park, 2000 (14pp)
Smith, D.W. et al. Yellowstone Wolf Project: Annual Report 1999. National Park Service, Yellowstone Center for Resources, Yellowstone National Park, 1999.
Smith, D. W. et al. Yellowstone Wolf Project: Annual Report 1998. National Park Service, Yellowstone Center for Resources, Yellowstone National Park, 1998 (14pp)
Smith, M.E. et al. Review of Methods to Reduce Livestock Depradation: I. Guardian Animals. Acta Agriculturae Scandinavica, 50/4, November 2000, 279 (12pp).
Smith, R.P. Dark Days for the Isle Royale Wolves. Wisconsin Sportsman, 11/6, 1982, 54-59.
Soussan, T. 4 Wolves Back in Captivity. Albuquerque (NM) Journal, 14 June 2001.
Soussan, T. Trapped by Doubts. Albuquerque (NM) Journal, 31 August 2003, B1.
Soussan, T. Wolves Killed Catron Calves. Albuquerque (NM) Journal, 2 April 2002, A1.
Spangler, J. Big, Bad Wolf May Be In Utah. (Salt Lake City) Deseret News, 13 November 2002. A01.
Spangler, J. USU Wolf Report Provokes Growls. (Salt Lake City) Deseret News, 8 December 2002, A01.
Spaulding, R. et al. Observer Bias and Analysis of Gray Wolf Diets from Scat. Wildlife Society Bulletin, 28/4, Winter 2000, 947.
Speer, O.H. A Foot in the Trap, A Head in the Snare. The Alaska Sportsman, 10/1, January 1944, 12-13, 28-29.
Speer, O.H. Wise Old Wolves. The Alaska Sportsman, 5/2, February 1939, 25-27.
Spence, C.E. et al. Surgical Sterilization of Free-Ranging Wolves. Canadian Veterinary Journal, 40, 118-121.
Stahler, D.R. et al. The Acceptance of a New Breeding Male Into a Wild Wolf Pack. Canadian Journal of Zoology, 80, February 2000, 360-365.
Stahler, D.R. et al. Common Ravens...Preferentially Associate with Gray a Foraging Strategy in Winter. Animal Behaviour, 64, August 2002, 283.
Stahler, D.R. Interspecific Interactions Between the Common Raven (Corvus corax) and the Gray Wolf (Canis lupus) in Yellowstone National Park, Wyoming: Investigations of A Predator and Scavenger Relationship. Unpublished thesis, 2001, University of Vermont.
Stahler, D.R. et al. Yellowstone Wolves and GPS Collars: New Insights from New Technology. World Wolf Congress 2003.
Stanwell-Fletcher, J.F. Three Years in the Wolves' Wilderness. Natural History, 49/3, March 1942, 137-147.
Staff. Ontario Announces Strategy to Protect Algonquin Wolves. Canada NewsWire, 6 November 2001.
Staff. The Wolf Question: Should Wolves Be Reintroduced in the Northeast...? Portland (MA) Press Herald, 28 October 2001, 1K.
Stark, Mike. Officials Continue to Probe 5 Wolf Deaths. Billings Gazette, 10 December 2003.
Stark, Mike. Wolf Plans Get Experts' Approval. Billings Gazette, 2 December 2003.
Stebler, A.M. The Status of the Wolf in Michigan. Journal of Mammalogy, 25/1, 1944, 37-43.
Stein, T. Colorado Soon to Have Wolves at the Door. The Denver Post, 18 May 2003, A-01.
Stein, T. Rockies Ranchers Adapt to Reality of Wolves. Denver Post, 8 July 2001.
Stein, T. Wolf Killing Probed: Poisonings Kill At Least 4 Animals in Idaho. Denver Post, 1 August 2001.
Stein, T. Wolves Near Success Status: Packs Grow but Hurdles Linger. Denver Post, 20 September 2001.
Stein, T. Wolves' Return Faces a Rocky Landscape. Denver Post, 9 July 2001.
Stein, T. Yellowstone Packs Key to Diversity. Denver Post, 9 July 2001.
Stenlund, M.H. A Field Study of the Timber Wolf (Canis lupus) on the Superior National Forest, Minnesota. Minn. Dep. Conserv. Technical Bulletin 4, 1955 (55pp).
Stenlund, M.H. Our Timber Wolves. Outdoor America, April 1966, 14-16.
Stenlund, M.H. Timber Wolf: Saint or Sinner! Minnesota Naturalist, 3/2, December 1952.
Stepanov, Y.V. & S.B. Pole. Numbers of Wolves and the Attitude Towards Them in Kazahkstan During Recent Decades. Journal of Wildlife Resources, 1/3, 1996, 321-322.
Stephens, P. W. & R. O. Peterson. Wolf Avoidance Strategies of Moose. Holarctic Ecology 7, 1987, 239-244.
Stephenson, R.O. Characteristics of Exploited Wolf Populations. (Final Report, Federal Aid in Wildlife Restoration), Alaska Department of Fish and Game, 1978.
Stephenson, R.O. Characteristics of Wolf Den Sites. Alaska Board of Fish & Game, Proj. W-17-R-06,  1974.
Stephenson, R.O., et al. Wolf Biology and Management in Alaska--Update to 1991. In (Carbyn, L.N., et al., eds.) Ecology and Management of Wolves in a Changing World. Canadian Circumpolar Institute, 1995.
Stephenson, R.O. & R.T. Ahgook. The Eskimo Hunter's View of Wolf Ecology and Behavior. In (Fox, M.W., ed.) The Wild Canids: Their Systematics, Behavioral Ecology, and Evolution, 1975, 286-291.
Stephenson, R.O. et al. Wolf Biology and Management in Alaska, 1981-1992. In (Carbyn, L.N. et al., eds.) Ecology and Conservation of Wolves in a Changing World, Canadian Circumpolar Institute, 1995, 43-54.
Stephenson, R.O. & D. ames. Wolf Movements and Food Habits in Northwest Alaska. In (Harrington, F.H. & P.C. Paquet, eds.) Wolves of the World, 1982, 26-42.
Steyn, Mark. Predators Aren't Looking for Peace. Chicago Sun-Times, 7 September 2003.
Stoehr, K.S. (ed). The Reintroduction of Wolves to Yellowstone and Central Idaho. International Wolf, 5/2, Summer 1995, 1-32.
Strickland, D. Wolf Howling in Parks: The Algonquin Experience in Interpretation. In (Carbyn, L.N., ed.) Wolves in Canada and Alaska: Their Status, Biology, and Management, Canadian Wildlife Service Report Series, Number 45, 1983, 93-95.
Sumanik, R.S. Wolf Ecology in the Kluane Region, Yukon Territory. M.Sc. thesis, Michigan Technical University, 1987.
Sundquist, A.K. et al. Y Chromosome Haplotyping in Scandinavian Wolves Based on Microsatellite Markers. Molecular Ecology, 10/8, 2001, 1959-1966.
Switalski, T. Coyote Foraging Ecology and Vigilance in Response to Gray Wolf Reintroduction in Yellowstone National Park. Canadian Journal of Zoology, 81, June 2003, 985-993.
Switalski, T.A. et al. Wolves in Utah: An Analysis of Potential Impact and Recommendations for Management. Utah Cooperative Fish and Wildlife Research Unit, Utah State University, Environmental Issue, 10, 2002.